Urocyon (Greek: "tailed dog"[ 7] ) is a genus of Canidae which includes the gray fox (Urocyon cinereoargenteus ) and the island fox (Urocyon littoralis ).[ 2] These two fox species are found in the Western Hemisphere. Whole genome sequencing indicates that, among living canids, Urocyon is sister to the remaining genera.[ 8] Fossils of what is believed to be the ancestor of the gray fox, Urocyon progressus , have been found in Kansas and date to the Upper Pliocene ,[ 9] with some undescribed specimens dating even older.[ 10]
Extant species
Genus Urocyon – Baird, 1857 – two species
Common name
Scientific name and subspecies
Range
IUCN status and estimated population
Gray fox
Urocyon cinereoargenteus (Schreber, 1775)
Southern half of North America from southern Canada to the northern part of South America (Venezuela and Colombia), excluding the mountains of northwestern United States
LC
Island fox
Urocyon littoralis (Baird, 1857)
U. l. littoralis (the nominate subspecies ) of San Miguel Island,
U. l. dickeyi of San Nicolas Island,
U. l. catalinae of Santa Catalina Island,
U. l. clementae of San Clemente Island,
U. l. santacruzae of Santa Cruz Island, and
U. l. santarosae of Santa Rosa Island.
Channel Islands (off the coast of Southern California)
NT
Extinct species
Skull cast of U. minicephalus , Florida Museum of Natural History
Species name
Type specimen era and location[ 4]
U. citrinus
Tedford et al. , 2009. Early Irvingtonian, Citrus County, Florida .
U. galushai
Tedford et al. , 2009. Late Blancan, San Simon Valley, Graham County, Arizona.
U. minicephalus
Martin, 1974. Late Irvingtonian, Sumter County, Florida.
U. progressus
Stevens, 1965. Early Blancan, Meade County, Kansas. A later review found that the material represents Urocyon , but because of its fragmentary nature cannot be diagnosed as to species.[ 4]
U. webbi
Tedford et al. , 2009. Middle Hemphillian, Citrus County, Florida.
Cozumel fox
The Cozumel fox is a critically endangered small gray fox found on the island of Cozumel, Mexico . The last reported sighting was in 2001, but surveys focusing on this species have not yet been carried out.[ 11] In September 2023, a live Cozumel fox was rescued from a road and subsequently released.[ 12] [ 13]
The Cozumel fox has not been scientifically described, but is a dwarf form, like the island fox. Compared to the island fox, however, it is slightly larger, being up to three-quarters the size of the gray fox. No skins or complete skulls of the Cozumel fox exist in any museum exhibitions, so scientists have mainly examined sub-fossils collected during archaeological excavations of Mayan civilizations who inhabited the island about 1,500–500 years ago. Upon evaluating bones from about 12 adult individuals, scientists have concluded that the Cozumel fox is extremely small – approximately 60-80% the body size of other mainland specimens.[ 14]
The fox had been isolated on Cozumel for at least 5,000 years, and probably far longer. These dates would indicate that Urocyon had colonized the island before the first arrival of humans there.[ 14]
References
^ "Fossilworks: Urocyon" .
^ a b Wozencraft, W. C. (2005). "Order Carnivora" . In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 582– 583. ISBN 978-0-8018-8221-0 . OCLC 62265494 .
^ Baird, S. 1857. Reports of explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean. vol.8(1):121, 138. Vulpinae
^ a b c d e Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic systematics of the North American fossil Caninae (Carnivora: Canidae )" (PDF) . Bulletin of the American Museum of Natural History . 325 : 1– 218. doi :10.1206/574.1 . hdl :2246/5999 . S2CID 83594819 .
^ Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press. ISBN 978-0-8018-8221-0 . OCLC 62265494 .
^ Schreber, Johann Christian Daniel; Goldfuss, Georg August; Wagner, Andreas Johann (1774). Die Säugthiere in Abbildungen nach der Natur, mit Beschreibungen . Vol. 3. p. 361. doi :10.5962/bhl.title.67399 . (
The mammals in illustrations according to nature, with descriptions)
^ Urocyon.
^ Lindblad-Toh, K.; Wade, C.M.; Mikkelsen, T.S.; Karlsson, E.K.; Jaffe, D.B.; Kamal, M.; et al. (2005). "Genome sequence, comparative analysis and haplotype structure of the domestic dog" . Nature . 438 (7069): 803– 819. Bibcode:2005Natur.438..803L . doi :10.1038/nature04338 . PMID 16341006 .
^ Stevens, M.S. (1965). "A new species of Urocyon from the Upper Pliocene of Kansas". Journal of Mammalogy . 46 (2): 265– 269. doi :10.2307/1377846 . JSTOR 1377846 .
^ Prevosti, F.J.; Rincóon, A.D. (2007). "A new fossil canid assemblage from the late Pleistocene of northern South America: The canids of the Inciarte asphalt pit (Zulia, Venezuela), fossil record and biogeography". J. Paleontol. 81 (5): 1053– 1065. doi :10.1666/pleo05-143.1 . hdl :11336/242701 . S2CID 131259363 .
^ Cuarón, Alfredo D.; Martinez-Morales, Miguel Angel; McFadden, Katherine W.; Valenzuela, David & Gompper, Matthew E. (2004). "The status of dwarf carnivores on Cozumel Island, Mexico". Biodiversity and Conservation . 13 (2): 317– 331. CiteSeerX 10.1.1.511.2040 . doi :10.1023/B:BIOC.0000006501.80472.cc . S2CID 25730672 .
^ Maya, Riviera (2023-09-18). "Rarely seen gray fox rescued 'disoriented' from Cozumel highway" . Riviera Maya News . Retrieved 2024-12-09 .
^ Wilkinson, Laura (2023-09-21). "Cozumel Wildlife Grey Fox - Cozumel 4 You" . Retrieved 2024-12-09 .
^ a b Gompper, M.E.; Petrites, A.E.; Lyman, R.L. (2006). "Cozumel Island fox (Urocyon sp.) dwarfism and possible divergence history based on subfossil bones" . Journal of Zoology . 270 (1): 72– 77. doi :10.1111/j.1469-7998.2006.00119.x . ISSN 1469-7998 .
Feliformia ("cat-like" carnivorans)
Feliformia
African palm civet (Nandinia binotata )
Feloidea
Prionodon (Asiatic linsangs)
Banded linsang (P. linsang )
Spotted linsang (P. pardicolor )
Felinae sensu stricto
Bay catlineage
Caracallineage
Leopardus Lynx Pumalineage
Leopard catlineage
Felis
Viverroidea
Viverroidea
Viverridae
Palm civets
Hemigalinae
Owston's palm civet (Chrotogale owstoni )
Otter civet (Cynogale bennettii )
Hose's palm civet (Diplogale hosei )
Banded palm civet (Hemigalus derbyanus )
Paradoxurinae
Binturong (Arctictis binturong )
Small-toothed palm civet (Arctogalidia trivirgata )
Sulawesi palm civet (Macrogalidia musschenbroekii )
Masked palm civet (Paguma larvata )
Paradoxurus
Viverrinae sensu lato
Viverrinaesensu stricto
Small Indian civet (Viverricula indica )
African civet (Civettictis civetta )
Viverra
Malabar large-spotted civet (V. civettina )
Large-spotted civet (V. megaspila )
Malayan civet (V. tangalunga )
Large Indian civet (V. zibetha )
Genettinae
Poiana(African linsangs)
Central African oyan (P. richardsonii )
West African oyan (P. leightoni )
Genetta(genets)
Abyssinian genet (G. abyssinica )
Angolan genet (G. angolensis )
Bourlon's genet (G. bourloni )
Crested servaline genet (G. cristata )
South African small-spotted genet (G. felina )
Common genet (G. genetta )
Johnston's genet (G. johnstoni )
Letaba genet (G. letabae )
Rusty-spotted genet (G. maculata )
Pardine genet (G. pardina )
Aquatic genet (G. piscivora )
King genet (G. poensis )
Schouteden's genet (G. schoutedeni )
Servaline genet (G. servalina )
Hausa genet (G. thierryi )
Cape genet (G. tigrina )
Giant forest genet (G. victoriae )
Herpestoidea
Eupleridae(Malagasy carnivorans)
Euplerinae(Malagasy civets)
Eupleres (falanoucs)
Eastern falanouc (E. goudotii )
Western falanouc (E. major )
Galidiinae(vontsira)
Ring-tailed vontsira (Galidia elegans )
Galidictis
Broad-striped Malagasy mongoose (G. fasciata )
Grandidier's mongoose (G. grandidieri )
Narrow-striped mongoose (Mungotictis decemlineata )
Salanoia
Suricata
Liberian mongoose (Liberiictus kuhni )
Mungos
Gambian mongoose (M. gambianus )
Banded mongoose (M. mungo )
Pousargues's mongoose (Dologale dybowskii )
Helogale
Ethiopian dwarf mongoose (H. hirtula )
Common dwarf mongoose (H. parvula )
Crossarchus(kusimanses)
Alexander's kusimanse (C. alexandri )
Angolan kusimanse (C. ansorgei )
Common kusimanse (C. obscurus )
Flat-headed kusimanse (C. platycephalus )
Marsh mongoose (Atilax paludinosus )
Long-nosed mongoose (Xenogale naso )
Urva(Asian mongooses)
Small Indian mongoose (U. auropunctata )
Short-tailed mongoose (U. brachyura )
Indian grey mongoose (U. edwardsii )
Indian brown mongoose (U. fusca )
Javan mongoose (U. javanica )
Collared mongoose (U. semitorquata )
Ruddy mongoose (U. smithii )
Crab-eating mongoose (U. urva )
Stripe-necked mongoose (U. vitticolla )
White-tailed mongoose (Ichneumia albicauda )
Yellow mongoose (Cynictis penicillata )
Selous's mongoose (Paracynictis selousi )
Meller's mongoose (Rhynchogale melleri )
Bdeogale
Bushy-tailed mongoose (B. crassicauda )
Jackson's mongoose (B. jacksoni )
Black-footed mongoose (B. nigripes )
Herpestes(slender mongooses)
Angolan slender mongoose (H. flavescens )
Egyptian mongoose (H. ichneumon )
Somalian slender mongoose (H. ochracea )
Cape gray mongoose (H. pulverulenta )
Common slender mongoose (H. sanguinea )
Vulpini
Nyctereutes(raccoon dogs) Vulpes (true foxes )
Canini (true dogs)
Cerdocyonina(zorro)
Speothos
Short-eared dog (Atelocynus microtis )
Crab-eating fox (Cerdocyon thous )
Lycalopex(South American foxes)
Canina(wolf-like canids)
Arctoidea
Ailuropoda Tremarctos Ursinae
Otariidae (eared seals)
Callorhinus (northern fur seals)
Northern fur seal (C. ursinus )
Otariinae (sea lions )
Neophoca
Australian sea lion (N. cinerea )
New Zealand sea lion (Phocarctos hookeri )
Arctocephalus (southern fur seals)
South American fur seal (A. australis )
Australasian fur seal (A. forsteri )
Galápagos fur seal (A. galapagoensis )
Antarctic fur seal (A. gazella )
Juan Fernández fur seal (A. philippii )
Brown fur seal (A. pusillus )
Guadalupe fur seal (A. townsendi )
Subantarctic fur seal (A. tropicalis )
Phocinae("northern seals")
Bearded seal (Erignathus barbatus )
Hooded seal (Cystophora cristata )
Phocini
Monachinae("southern seals")
Musteloidea
Mephitidae (skunks)
Conepatus(hog-nosed skunks)
Molina's hog-nosed skunk (C. chinga )
Humboldt's hog-nosed skunk (C. humboldtii )
American hog-nosed skunk (C. leuconotus )
Striped hog-nosed skunk (C. semistriatus )
Mephitis Mydaus(stink badgers)
Sunda stink badger (M. javanensis )
Palawan stink badger (M. marchei )
Spilogale(spotted skunks)
Southern spotted skunk (S. angustifrons )
Western spotted skunk (S. gracilis )
Eastern spotted skunk (S. putorius )
Pygmy spotted skunk (S. pygmaea )
Procyonidae
Bassariscus
Ringtail (B. astutus )
Cacomistle (B. sumichrasti )
Procyon(raccoons) Bassaricyon(olingos)
Eastern lowland olingo (B. alleni )
Northern olingo (B. gabbii )
Western lowland olingo (B. medius )
Olinguito (B. neblina )
Nasuina(coatis)
Nasua Nasuella(mountain coatis)
Eastern mountain coati (N. meridensis )
Western mountain coati (N. olivacea )
Mustelidae
Mustelidae
Mellivora
Arctonyx(hog badgers)
Northern hog badger (A. albogularis )
Greater hog badger (A. collaris )
Sumatran hog badger (A. hoevenii )
Meles(Eurasian badgers)
Melogale(ferret-badgers)
Vietnam ferret-badger (M. cucphuongensis )
Bornean ferret badger (M. everetti )
Chinese ferret-badger (M. moschata )
Javan ferret-badger (M. orientalis )
Burmese ferret-badger (M. personata )
Formosan ferret-badger (M. subaurantiaca )
Guloninae
Ictonychinae
Lyncodontini
Patagonian weasel (Lyncodon patagonicus )
Galictis(grisons)
Lesser grison (G. cuja )
Greater grison (G. vittata )
Ictonychini(African polecats)
Vormela
Marbled polecat (V. peregusna )
African striped weasel (Poecilogale albinucha )
Ictonyx
Saharan striped polecat (I. libycus )
Striped polecat (I. striatus )
Lontra Enhydra
Spotted-necked otter (Hydrictis maculicollis )
Lutra Lutrogale
Smooth-coated otter (L. perspicillata )
Aonyx
African clawless otter (A. capensis )
Asian small-clawed otter (A. cinereus )
Congo clawless otter (A. congicus )
Mustelinae
Neogale(New World weasels)
subgenus Mustela (paraphyletic) subgenus Lutreola (paraphyletic)
Japanese weasel (M. itatsi )
European mink (M. lutreola )
Indonesian mountain weasel (M. lutreolina )
Malayan weasel (M. nudipes )
Siberian weasel (M. sibirica )
Back-striped weasel (M. strigidorsa )
subgenus Putorius
† Hesperocyoninae
Hesperocyon
Mesocyon
Cynodesmus
Sunkahetanka
Philotrox
Enhydrocyon
Paraenhydrocyon
Caedocyon
Ectopocynus
Osbornodon
O. brachypus
O. fricki
O. iamonensis
O. renjiei
O. sesnoni
O. scitulus
O. wangi
Archaeocyon
Oxetocyon
Otarocyon
Rhizocyon
Phlaocyonini
Phlaocyon
P. achoros
P. annectens
P. latidens
P. leucosteus
P. mariae
P. marslandensis
P. minor
P. multicuspus
P. taylori
P. yatkolai
Borophagini
Cormocyon
Desmocyon
Metatomarctus
Euoplocyon
Psalidocyon
Microtomarctus
Protomarctus
Tephrocyon
Cynarctina Aelurodontina Borophagina
Paratomarctus
Carpocyon
Protepicyon
Epicyon
Borophagus
B. diversidens
B. dudleyi
B. hilli
B. littoralis
B. orc
B. parvus
B. pugnator
B. secundus
Caninae
Caninae
Vulpini
Nyctereutes (raccoon dogs)
† N. abdeslami
† N. donnezani
† N. lockwoodi
† N. megamastoides
† N. sinensis
† N. tingi
† N. vinetorum
Vulpes (true foxes)
† V. hassani
† V. odessana
† V. praeglacialis
† V. qiuzhudingi
† V. riffautae
† V. rooki
† V. skinneri
† V. stenognathus
Canini (true dogs)
Cerdocyonina(zorro)
Speothos
† Pleistocene bush dog (S. pacivorus )
† Dusicyon
Falkland Islands wolf (D. australis )
D. avus
D. cultridens
Lycalopex(South American foxes)
† L. cultridens (?)
† L. ensenadensis
Canina (wolf-like canids)
Canina (wolf-like canids)
Canina
† Dire wolf (Aenocyon )
† Sardinian dhole (Cynotherium )
† Trinil dog (Mececyon )
† Merriam's dog (Megacyon )
† Xenocyon
† Eucyon
E. davisi
E. debonisi
E. ferox
E. intrepidus
E. khoikhoi
E. kuta
E. marinae
E. minor
E. monticinensis
E. skinneri
E. wokari
E. zhoui
Lycaon Dhole (Cuon alpinus )
† C. adoxus
† C. antonii
† C. apolloniensis
† Armbruster's wolf (C. armbrusteri )
† C. arnensis
† C. borjgali
† C. cedazoensis
† C. chihliensis
† Edward's wolf (C. edwardii )
† Etruscan wolf (C. etruscus )
† C. lepophagus
† Mosbach wolf (C. mosbachensis )
† C. nehringi
† C. palmidens
† C. variabilis
† Prehistoric† Recently extinct
Sicilian wolf (C. l. cristaldii )
Hokkaido wolf (C. l. hattai )
Japanese wolf (C. l. hodophilax )
Austro-Hungarian wolf
Synonymized under C. l. arctos : Bernard's wolf (C. l. bernardi )
Synonymized under C. l. occidentalis : Kenai Peninsula wolf (C. l. alces )
Manitoba wolf (C. l. griseoalbus )
Synonymized under C. l. nubilus : Newfoundland wolf (C. l. beothucus )
Cascade mountain wolf (C. l. fuscus )
Mogollon mountain wolf (C. l. mogollonensis )
Texas wolf (C. l. monstrabilis )
Southern Rocky Mountain wolf (C. l. youngi )
See also : Evolution of the wolf
Category:Extinct canids
Authority control databases
National Other